Anomalocaris

Jak číst taxoboxAnomalocaris
Stratigrafický výskyt: Střední kambrium, asi před 516 až 505 miliony let
alternativní popis obrázku chybí
kresba anomalocarise
Vědecká klasifikace
Říšeživočichové (Animalia)
OdděleníEcdysozoa
Kmenčlenovci (Arthropoda)
TřídaDinocarida
ŘádRadiodonta
ČeleďAnomalocaridae
RodAnomalocaris
Whiteaves, 1892
Některá data mohou pocházet z datové položky.

Anomalocaris byl asi 60 až 90 (nebo dokonce 200) cm dlouhý dravý bezobratlý živočich (členovec), žijící v období prvohorního kambria. Lovil pravděpodobně drobné členovce, jako byli trilobiti. Byl to jeden z prvních velkých dravců této planety. Měl dvě dlouhá chapadla, kterými si podával kořist k ústům, ve kterých měl zuby. Anomalocaris měl dvě poměrně velké oči. Fosilní pozůstatky tohoto tvora byly objeveny v burgeské vrstvě v Kanadě, ale také v Číně, Austrálii, Utahu a v dalších státech.[1] Plaval pomocí zvlněných výstupků, které mu vybočovaly z těla. Anomalocaris dokázal rychle a obratně plavat a vyhledávat kořist, kterou svými chapadly a zuby lovil.

V populární kultuře

Anomalocaris se objevil v množství vědeckých filmových dokumentů. V dvojdílném dokumentu První život od Davida Attenborougha byl v animaci podle pravdy popisován jako jeden z prvních velkých dravců, ukázány byly rovněž zkameněliny rodu. Dále se objevil v prvním díle dokumentu Triumf obratlovců od Davida Attenborougha, kde figuruje na začátku dokumentu v animaci, jak pronásleduje myllokunmingiu, vyskytl se i v titulu Země: vznik planety a první části trojdílného seriálu Putování s pravěkými monstry od BBC.

Reference

  1. Hou, X.-G.; Bergström, J. and Ahlberg, P. (1995). Anomalocaris and other large animals in the Lower Cambrian Chengjiang fauna of southwest China. Geologiska Föreningens i Stockholm Forhandlingar, 117: 163-183.

Literatura

  • Whittington H. B.; Briggs, D. E. (1985). "The largest Cambrian animal, Anomalocaris, Burgess Shale, British Columbia". Philosophical Transactions of the Royal Society B. 309 (1141): 569–609. doi:10.1098/rstb.1985.0096

Externí odkazy

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Autor: , Licence: CC BY 2.5
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Peytoia nathorsti Laggania cambria oral cone, Burgess Shale.jpg
Autor: James St. John, Licence: CC BY 2.0

Peytoia nathorsti Walcott, 1911 (=Laggania cambria Walcott, 1911) mouthpiece (~5.25 x ~4.25 cm), preserved as a carbonized film in slightly metamorphosed shale from the Middle Cambrian-aged Burgess Shale of southwestern Canada (YPM 5825, Yale University’s Peabody Museum, New Haven, Connecticut, USA). This fossil was formerly identified as a fossil jellyfish (Animalia, Cnidaria, Scyphaozoa) and had been misidentified as the mouthpiece of Anomalocaris canadensis Whiteaves, 1892 until the restudy done by Daley & Bergström 2012.


The Middle Cambrian-aged Burgess Shale is the most famous fossil deposit on Earth. It is located near the town of Field in Yoho National Park, southeastern British Columbia, western Canada. The deposit is famous for its spectacular soft-bodied preservation - the organisms have had their appendages & internal organs preserved. Many tens of thousands of fossils have been collected from the Burgess Shale Formation over the last century. Including known, but unnamed species, and excluding known or demonstrable junior synonyms, the Burgess Shale biota totals at least ~280 species.

Many claim that Charles Walcott discovered the Burgess Shale Lagerstätte (as soft-bodied fossil deposits are called by paleontologists) in 1909. However, it was actually discovered in 1886 or 1888 by Richard McConnell, based on anomalocarid appendage material from Mt. Stephen, in the Campsite Cliff Member of the Burgess Shale Formation. The main collecting localities have been two quarries (Walcott Quarry & Raymond Quarry) on the western side of the ridge connecting Mt. Field and Wapta Mountain a little north-northeast of Field. Numerous other smaller localities have been identified in the same area & for many, many kilometers to the south. Collecting at the Burgess Shale was most intense in 1910-1917 (Charles Walcott), 1925-1930 (Harvard’s Museum of Comparative Zoology), 1966-1967 (Geological Survey of Canada), and 1975-2000s (Royal Ontario Museum).

Some of the most celebrated fossil organisms from the Burgess Shale are the radiodonts. These creatures had medium-sized to large bodies (ranging from <10cm to anout half of a meter long) with a head having a pair of frontal appendages & short-stalked eyes & a pineapple ring-shaped mouth, plus a body with two lateral rows of swimming flaps.

The first radiodont fossils (isolated frontal appendages) were discovered in the 1880s from British Columbia’s Burgess Shale Formation. They were identified as shrimp bodies lacking heads (see photo elsewhere in this album).

In general, paleontologists didn’t recognize that these Anomalocaris fossils represented parts of a much larger organism. The true nature of the complete Anomalocaris organism wasn’t realized until very rare complete specimens were excavated from Burgess Shale quarries by the Royal Ontario Museum.

The mouthpiece of Peytoia nathorsti turned out to be an already-known fossil previously identified as a jellyfish. Peytoia mouthpieces resemble pineapple rings, consisting of a radially arranged series of plates with inward-directed spines. Some bitemarks found on trilobites & other fossil arthropods are consistent with radiodont mouthpiece morphology.

The unusual body plan of radiodont does not fit into any traditional arthropod group, so new high-level taxa have been created to accomodate it (see Collins, 1996, Journal of Paleontology 70: 280-293).

Classification: Arthropoda, Dinocaridida, Radiodonta, Hurdiidae

Stratigraphy: Walcott Quarry Member, Burgess Shale Formation, Ptychagnostus praecurrens Interval-zone, lower Marjuman Stage, middle Middle Cambrian.

Locality: Walcott Quarry, western side of ridge between Mt. Field & Wapta Mountain, north-northeast of the town of Field, southeastern British Columbia, southwestern Canada.
Anomalocaris BW.jpg
Autor: Nobu Tamura, Licence: CC BY 3.0
Anomalocaris canadensis, the top predator from the middle Cambrian Burgess Shale of British Columbia, pencil drawing, digital coloring.
Anomalocaris Mt. Stephen.jpg
Anomalocaris "arm" from the Mt. Stephen Trilobite Beds, Middle Cambrian, near Field, British Columbia, Canada.